Monday, November 17, 2025

Insect Study Resources: Guides, Research, and Learning Tools from the Vermont Entomology Academy


The Vermont Entomology Academy is committed to making insect science accessible to learners of all backgrounds—from beginners discovering their first beetle to naturalists documenting new state records. To support that mission, we’ve curated a streamlined reference list of the most reliable field guides, research papers, articles, online courses, videos, and podcasts, organized by the major insect orders.

Whether you’re identifying backyard species, preparing for fieldwork, or deepening your understanding of insect ecology, this collection provides trusted resources across the full spectrum of entomology. Explore, learn, and follow your curiosity—one order at a time. 

You do not need to know the science of insects to be amazed by the insects. But we can be inspired even more when we learn a little about them.


Vermont Entomology Recommended Resources 



Family or Description

Link

All Insects 




Atlas of Life

VCE Wildlife Atlases *various


Atlas of Life

VCE: The Vermont Atlas of Life ***


Checklist

VCE: Checklist of Vermont Species


Guides

Agriculture and Agri-Food Entomological Monographs


Guides

BugFinder


Guides

BugGuide. ***


Guides

Canadian Journal of Arthropod Id


Guides

Discover Life ***


Guides

Insect Identification – Online Resources


Guides

US and Canada Bug Identification Guides


Guides to British insects.

Mike’s Insect keys.  Dichotomous keys are designed for use on screen.


Photos

Insect Photo Collection on Flickr


Species Profiles

Explore Species Profiles


VT Insects

Est. 21.4k insect species in VT

Collembola




Springtails

The Collembola of North America, North of the Rio Grande: A Taxonomic Analysis - 4 Volumes / grinnell-32544.pdf

Coleoptera: Beetles.




Beetles

VCE: Tiger Beetles of Vermont


Beetles

VCE: Vermont Lady Beetle Species Profiles


Fireflies

13 species known in VT


Fireflies

Field Guides and ID Keys


Fireflies

Firefly Anatomy and Flash Patterns


Fireflies

Firefly Photography Tips


Fireflies

Firefly Photography Tips II


Fireflies

Firefly Photography Tips III and IV


Fireflies

Firefly Survey


Fireflies

iNat link


Fireflies

Preliminary List


VT Beetles

Over 1k species in VT. One out of every four creatures on the planet is a beetle. 

Diptera: Flies




Craneflies

Crane flies of eastern North America


Craneflies

The Craneflies (Diptera: Tipulidae) of Pennsylvania


Symphyta / Sawflies

Electronic World Catalog of Symphyta - symphyta on iNaturalist


Syrphidae

Bee flies (Bombyliidae)


Syrphidae

Field/Photo ID for Flies


Syrphidae

Key to the Syrphus of North America


Tachinid Flies Family Tachinidae

Taxonomic and Host Catalogue of the Tachinidae of America North of Mexico

Diptera: Fly Genus detail




Condylostylus

Complex Condylostylus sipho: Yellow femora and tibiae combined with marked wings are indicators for the sipho group, rather than in patibulatus, which also has marked wings, but which has all-black legs in both sexes. Sadly, so far, females are indistinguishable between species in this group. Very good side shots of the legs are best for identifying sipho males to species. - alisonnetta on iNaturalist


Cylindromyia

There are 17 species of Cylindromyia in the Nearctic (http://www.nadsdiptera.org/Tach/Nearctic/CatNAmer/Genera/Cylindromyia.html). Cylindromyia bicolor is not one of them (it is a European species). The ID engine of iNaturalist is misleading people to put a species name on their Cylindromyia when they should not. - jhskevington, iNaturalist


Merodontini

Common Merodontini of the Northeast by Even and Zachary Dankowicz. “Merodontini”: A tribe of hoverflies, also known as "bulb flies," within the family Syrphidae. 


Orthorrhapha:Family   Rhagionidae

This species has a white frons (seen in females only); crescents or complete bands on the first few abdominal segments; dark stigmata in the wings; and often near-white fore tibiae. There's also a table of information on them in the Fly Guides under Orthorrhapha. Table of information on Rhagio in the Fly Guides (search site using 'Rhagionidae', and look in the 'Notes' section). Alisonnetta iNaturalist 

Hemiptera: True bugs




VT True Bugs

True Bugs: At least 80 species in VT

Hymenoptera: Ants, bees, and wasps




CAUTION on making IDs in  iNaturalist 

When making an ID on iNaturalistplease leave species ID to the experts unless you are very sure and can recognize the features that make it that specific species. Many species cannot be identified from a photo. Many species are very similar to other species. IF in doubt, ID it at a higher rank (taxon). This approach helps maintain data integrity. ID at the taxon rank (level), you are sure of. You can write “I think it is this (____) species” as a comment, even if the (menu selection) is more general.                                                                                                   ReCap: If you are only sure that it is an insect, ID it as an insect. If you are sure it is a bee, but not sure of a more detailed taxon for the bee, ID it as a Bee. If you are sure it is Genus Bombus (Bumblebee), ID as Bombus. If you know the subgenus, go with that. But only ID at species if you can support the ID with anatomy details that make it that species.                                                                                                 Even a relatively few wrong and speculative IDs (particularly at the species level) ruin credibility. CAUTION: The Artificial Intelligence on iNaturalist makes ID recommendations, but they are often incorrect. 


Andrena

Andrena key


Atlas of Life

VCE: The Vermont Atlas of Life ***


Bee Anatomy

Bee Anatomy


Bees

Bee Photos. USGS Bee Lab


Bees

Bee species by percentage Chart


Bees

Bee Watching’s Field Guide to Wild Bees


Bees

Bees of Maryland: A Field Guide


Bees

Bees of NY and nesting information


Bees

Beyond the Honeybee: Wild Bees on the Vermont Landscape


Bees

How to make a habitat for ground nesting bees


Bees

Sharp-Eatman Guide - New York


Bees

State of Vermont's Wild Bees Conservation


Bees

State of Vermont's Wild Bees: Current Knowledge


Bees

State of Vermont’s Wild Bees 2022


Bees

The Bees of North Carolina - An Identification Guide


Bees

The Melissodes Project


Bees

VCE: Masked Bees (Genus Hylaeus)


Bees

VCE: The Bees of VT: Id by Species, Genera, Specialists by host plant or by habitat.


Bees

VCE: Vermont Bee Species


Bees

VCE: Vermont Wild Bee Guide


Bees 

351 known bee species in VT


Bees 

Bees of Canada


Bees 

VCE: Bees of VT Supplement Species Account (2025)


Bees & Wasps

Identifying Bees and Wasps


Bees and Plants

Pollen Specialist Native Bees: Native Plants Are Key To Their Survival - Heather Holm


Bees: Bumblebees

Bumblebees by Neylon


Bees: Green Bees

Metallic Green Sweat Bees (Bees of Minnesota)


Bees: Halictid (Green Bees)

North American Halictid Green Bee Genera Quick Guide, Clare Maffei


Bees: Specialist Bees

Pollen Specialist Bees of the Eastern United States, Jarrod Fowler, Sam Droege


Bumble Bees

17 Bumble Bee Species known in Vermont (4 have are no longer seen in VT)


Bumble Bees

Aberrant bee color patterns - iNaturalist project page


Bumble Bees

Bumble Bee Anatomy


Bumble Bees

Bumble Bee Field Guide


Bumble Bees

Bumble Bee ID By Illustrations (click on images to enlarge)


Bumble Bees

Bumble Bee Identification Guides Links


Bumble Bees

Bumble Bee Life Cycle: Graphic Art 


Bumble Bees

Bumble Bees of Illinois


Bumble Bees

Bumble Bees of the Eastern US


Bumble Bees

Comparing Bombus impatiens, bimaculatus, & griseocollis by neylon on inaturalist


Bumble Bees

Photos of Bumble Bees by Heather Helm


Bumble Bees

Resources to Boost your Bumble Bee Identification Skills


Bumble Bees

VCE: Bumble Bees (Genus Bombus)


Bumble Bees

VCE: Bumble Bees of New England


Bumble Bees

VCE: Bumble Bees of New England (illustrated guide)


Bumble Bees

VCE: Field Guide to the Bumble Bees of VT


Bumble Bees & Carpenter Bees

VCE: Bumble Bees and Carpenter Bees of Vermont (Rank)


Ichneumonid Wasp

Ichneumon wasp identification


Specialist Bees

Guide to Specialist Bees of Ohio and Floral Resources to Target


Specialist Bees

Pollen Specialist Bees of the Eastern United States and associated plants


Specialist Bees

VCE: Specialist Bees and associated plants


Terms that describe a bee

Glossary of Terms

Bees - Genus detail




A Genera List

VT Bee Genera (VCE)


A side by side: Green Sweat Bees

Metallic Green Sweat Bees


A Species List

BEE SPECIES OF VERMONT (VCE) 

Lepidoptera: Butterflies and Moths




Butterflies

31 common butterflies found in Vermont


Butterflies

Butterflies & Moths of North America


Butterflies

Butterfly Id


Butterflies

Catalogue of the Butterflies of the United States and Canada” by Jonathan P. Pelham


Butterflies

Identify a butterfly


Butterflies

Identifying Butterflies by Color


Butterflies

Sharon Wander’s Grass Skippers Tops Guide 


Butterflies

Wisconsin Butterflies


Moths

Mothing 101


Moths

Underwing Moths (Catocala) & Larvae A companion guide for iNaturalists bob_borth on iNaturalist


VT Butterflies

At least 120 Butterfly species in VT


VT Moths

2,051 species of moths are known from Vermont!

Odonata: Dragonflies and Damselflies




Dragonflies    

Vermont Damselfly and Dragonfly Atlas


VT Dragonflies    

101 Dragonflies and 45 Damselflies known in VT

Orthoptera: GrassGrasshoppers, Crickets and bush-crickets




Bush-crickets

Common Bush-crickets. British focus


Orthoptera

Orthoptera of Iowa


VT grasshoppers

 At least 91 species in VT

Misc Taxonomic Orders






Plants for insects




Plants

Pollinator Gardens – going beyond the plant lists to create robust habitat.

Field Guide Book




Ants

A Field Guide to the Ants of New England


Appalachian Brown vs Eyed Brown

Appalachian Brown vs Eyed Brown:Habitat: Is it within a wet woodland with sedges? (Appalachia). Is it confined to sedges in an open sunny field/marsh? (Eurydice). Both occur in edge habitat. Appy's are more likely to wander out of their habitat. Intermediate individuals may be difficult, if not impossible, to ID. Field marks vary; a combination of all characteristics should be considered. Color can be key to the species ID — best when seen in life vs photos/specimens. - hobiecat on iNaturalist


Aquatic insects

Turning Stones - Discovering the Life of Water


Bees

Bees of the Eastern United States Vol 1 and 2 by Theodore B. Mitchell


Bees

Common Bees of Eastern North America


Bees

The Bees in Your Backyard


Bees

The Solitary Bees


Beetles

Guide to Beetles


Beetles

Peterson Field Guides: Beetles


Bumble Bees

Bumble Bees of North America: An Identification Guide


Bumble Bees

Ohio State Univ. Links to resources 


Butterflies

A pocket guide to Butterflies and Moths


Butterflies

A Swift Guide to Butterflies of North America: Second Edition


Butterflies

Butterflies of Maine and the Canadian Maritime Provinces


Butterflies

Butterflies of the East Coast: An Observer's Guide 


Butterflies

Butterflies through Binoculars: The EastA Field Guide to the Butterflies of Eastern North America


Butterflies

Kaufman Field Guide to Butterflies of North America


Butterflies

Peterson Field Guide To Moths Of Northeastern North America


Caterpillars

Caterpillars of Eastern North America: A Guide to Identification and Natural History


Damselflies

Damselflies of the Northeast: A Guide to the Species of Eastern Canada & the Northeastern United States


Dragonflies & damselflies

A field guide to the dragonflies and damselflies of Massachusetts


Dragonflies & damselflies

Dragonflies and Damselflies of the East


Dragonflies & damselflies

Field Guide to The Dragonflies and Damselflies of Algonquin Provincial Park and the Surrounding Area


Fireflies

Fireflies, Glow-worms, and Lightning Bugs: Identification and Natural History of the Fireflies of the Eastern and Central United States and Canada


Fireflies

Flash Patterns of common fireflies 


Flies

Field Guide to the Flower Flies of Northeastern North America


Insects

Garden Insects


Insects

How to know the Immature Insects


Insects

Insects of New England and New York, Tom Murray


Insects

Insects: Their Natural History and Diversity: With a Photographic Guide to Insects of Eastern North Americansects-Their Natural History and Diversity- 2006. This is not a Field Guide in the usual sense but it is a gold mine of information, useful


Insects

Kaufman Field Guide to Insects of North America


Insects

The Peterson Field Guide Series, A Field Guide to the Insects of America North, Donald J. And White Borror (1970). Though obviously dated, I think it's the best general field guide 


Insects

What Insects Do, and Why


Lady Beetles

Are you a nine-spotted ladybug?


Lady Beetles

The Lady Beetles of Maine 


Tiger Beetles

A Field Guide to the Tiger Beetles of the United States and Canada


Wasps

The Social Wasps of North America

Other Insect Books




Animals

An Immense World


Ants (novel)

Anthill by E.O. Wilson


Bees

Dancing with Bees - a Journey back to Nature


Bees

The Mind Of A Bee


Beetles (fiction)

Beetle Boy


Bumble Bees

A Sting in the Tale


Bumble Bees

Bumble Bee Economics


Flies

The secret life of FLIES


Insects

Extraordinary Insects


Insects

Sex on Six Legs


Insects

The Thermal Warriors - Strategies of Insect Survival


Nature

Braiding Sweetgrass


Wasps

Wasps - The Astonishing Diversity of a Misunderstood Insect

Identification by Key




A1: Key Identification tools

Insect identification by key is a method using a dichotomous key, a tool with paired, contrasting choices (couplets) about physical traits (like wings, antennae, legs, body parts) that systematically narrow down possibilities until you identify the insect's order or species, guiding you down a "fork in the road" of characteristics until you reach a final answer. 


Andrena

Andrena key


Bees

Denny Johnson Agapostemon key


Bees: Halictid (Metallic Green bees)

Key to Metallic Green Halictid (Family Halictidae) Bees of Blandy Experimental Farm


Bees: Polistes ID key (2008)

Identification Atlas of the Vespidae (Hymenoptera, Aculeata) of the northeastern Nearctic region


Bees: Polistes ID key (2021)

Total-Evidence Phylogeny of the New World Polistes Lepeletier, 1836, Paper Wasps (Vespidae, Polistinae, Polistini)


Bees: Zadontomerus

(Small carpenter   bees)

Ohio Ceratina ID

Online Video Learning




Bees

My Garden of a Thousand Bees. Nature PBS


Bees

Sam Droege and Clare Maffei's how to 'bee' videos


Bees

The Unequal Cellophane Bee - Colletes inaequalis


Bees

US National Native Bee Monitoring RCN


Bees

Vermont's Wild Bees: Past, Present, and Future 10-5-23: Spence Hardy


Bumble Bees

Bumblebee Short Course For Community Scientists


Bumble Bees

Heather Holm, Bumble Bee Banquet: Selecting Native Plants for Bumble Bees


Bumble Bees

In the Life of the Bumble Bee Queen (and the cuckoo too). 


Bumble Bees

Keynote Speaker: Dr. Hollis Woodard, “The Ontogeny of Sociality in Bumble Bee Queens”


Bumble Bees

VCE: Bumble Bee Natural History and Conservation in Vermont


iNat download

How to download iNaturalist data onto your desktop (to a CSV.)


iNaturalist (value)

How Your iNaturalist Data Makes a Difference for Biodiversity


iNaturalist on a mobile device

Naturalist: uploading observations with mobile devices. Fewer functions than a desktop application. (How to video)


iNaturalist on Desktop

iNaturalist: uploading observations using desktops and laptops (How-to video)


iNaturalist use Tips

VCE: Lunchtime Learning Videos by Julia Pupko. Tips for using iNaturalist to it's fullest. (List of videos)


iNaturalist: Making IDs

Identifying on iNaturalist: How you can help! Using the Identity page starts at 18:43.


Insects

Bugs that Rule the World Nature PBS (requires PBS Subscription)


Insects (Film)

Beetle Queen Conquers Tokyo (2009). “Carefully look at nature, including insects. It’s actually a way of life.”


Phenology

Phenology in Focus: Exploring Plant Cycles with iNaturalist 


Specialist Bees

Bryan Danforth: Host-plant Specialist Bees – Biology, Biodiversity, Conserving Them in Your Backyard

Research Papers




BB’s and Flooding

Unveiling the submerged secrets: bumblebee queens' resilience to flooding


Bees

Zachary Portman’s list of Classic and helpful papers by Mitchener, Mitchell, and Jander


Bees - Augochloropsis

A review of the Augochloropsis (Hymenoptera, Halictidae) and keys to the shiny green Halictinae of the midwestern United States


Bees - Hymenoptera: Apoidea

An Annotated Checklist of the Bees (Hymenoptera: Apoidea) of Vermont with Conservation Status and Natural History Notes


Bees - Phenological trends

Ecological traits explain long-term phenological trends in solitary bees


Bees & Blueberries

UVM Graduate Research Feature: Leslie Spencer and Wild Pollinators


Bees at play

Do bumble bees play?


Bees behavior

Charles Henry Turner and the cognitive behavior of bees


Bees of Connecticut

Biodiversity of Bees (Hymenoptera: Apoidea: Anthophila) in Connecticut (USA), J. Asher, et al


Bees, nesting

Nest observations reveal hygienic nest architecture and behavior in an Arctic bumble bee species


Bees: Agapostemon

Taxonomy of Agapostemon angelicus and the A. texanus species complex (Hymenoptera, Halictidae) in the United States


Bees: Zadontomerus

(Small carpenter   bees)

Morphological and molecular delineation of a new species in the Ceratina dupla species-group (Hymenoptera: Apidae: Xylocopinae) of eastern North America


Bumble Bees

Simple and farmer-friendly bumblebee conservation: Straw bales as nest sites in agricultural landscapes


Climate Change: BB’s

Warming summer temperatures are rapidly restructuring North American bumble bee communities


Foraging range

Foraging ranges of solitary bees


iNaturalist

PDF: Community involvement in natural history: a growing opportunity for entomology and science


Insect pollinators

Insect pollinators: The time is now for identifying species of greatest conservation need - Hardy, McFarland et al


Lasioglossum & Agapostemon

Some relevant keys to look at for determining which characteristics should be seen for species ID: A review of the Augochloropsis (Hymenoptera, Halictidae) and keys to the shiny green Halictinae of the midwestern United States, AND Taxonomy of Agapostemon angelicus and the A. texanus species complex (Hymenoptera, Halictidae) in the United States AND Revision of the metallic Lasioglossum (Dialictus) of eastern North America (Hymenoptera: Halictidae: Halictini) - xianzx on iNaturalist


VT Atlas of Life

10th Anniversary Vermont Atlas of Life Report (2023)

Podcasts




Bees 

Vermont's bee populations are failing. Here's what you can do to help


Bumble Bees

Episode 19: What’s all the buzz about bumblebees? with Michael Whitehead

Articles




Bees

Bees are sentient’: inside the stunning brains of nature’s hardest workers


Bees

Bees may feel pain


Bees

If bumblebees can play, does it mean they have feelings? This study suggests yes


Bees

New Bees Discovered in Vermont with Worldwide Teamwork


Bees

New Vermont ‘Bee Team’ to Tackle Pollinator Threats


Bees

Nine new-to-Vermont bee species discovered - Hardy, Asher et al


Bees

Parasitized bees are self-medicating in the wild, study finds


Bees

Seeing the Forest for the Bees


Bees

Wild Bee Conservation


Bumble Bees

Conserving Bumblebees. Guidelines for creating and conserving habitat. 


Bumble Bees

Strategy To Protect State And Federally Recognized Bumble Bee Species Of Conservation Concern


Bumble Bees

Washington State Becomes First To Adopt A Statewide Strategy To Protect Bumble Bees


E-butterfly

e-Butterfly help center


iNaturalist (setting your license choice)

What are licenses? How can I update the licenses on my content?


iNaturalist Annotations

Annotate a Sighting: recommended fields for bees (when relevant) include “ Interaction->Visited flower of: “, “ Pollen present in corbicula or scopa”, and “ Interaction->Preyed upon by: ”


iNaturalist forum

iNat Forum Various themes or topics


iNaturalist primer

iNaturalist: How to guide from VCE


iNaturalist use Tips

How to use iNaturalist's search URLs (IN DEPTH)


iNaturalist use Tips

Tech Tip Tuesday. series of short articles. From little tricks that help you record better data to suggestions on how to make your observations easier to identify.


Odonata

Odonata Central User Guide


Odonata

Odonata Central: How to use the Odonata website.

Newsletters




Insects

Vermont Entomology Society Newsletters

iNaturalist Projects




Bees

Aberrant bee color patterns


Bees

Predation of Bees


Bees

Vermont Wild Bee Survey


Insects

Insects of Vermont


Lady Beetles

Vermont Lady Beetle Atlas


Mating bees

Mating Bees


Mimics

Bee and Wasp Mimics


Nesting bees

Nesting bees


Odonata

Odonata: Dragonflies and Damselflies


Plant/Pollinator

Pollinator Interactions on Plants (PIP) of the NE US


VT Stats by city

Vermont Biodiversity by city or town

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